BIOS101
General Biology

Prerequisites: Must Enroll in both BIOS 101 (lecture) and BIOS 101L (lab)

High school chemistry or equivalent strongly recommended. Not intended for most Life Sciences majors; such students should take LIFE 120-LIFE 120L and LIFE 121-LIFE 121L instead. This course does not count towards the biology major.

Description: Analysis of the structure, functions, and interactions of organisms from the molecular to the ecosystem levels. BIOS 101 provides the foundation for tomorrow's citizens to critically think about and evaluate issues in biology.

This course is a prerequisite for: ASCI 240; BIOC 321; BIOS 213; BIOS 213L; ENTO 308; FDST 401, FDST 801; PSYC 370

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 4 Science

Credit Hours:3

ACE:ACE 4 Science

BIOS101L
General Biology Laboratory

Prerequisites: Must enroll in both BIOS 101 (lecture) and BIOS 101L (lab)

Description: Laboratory exercises and experiments that complement material covered in BIOS 101.

This course is a prerequisite for: BIOC 321; BIOS 213; BIOS 213L; ENTO 308; PSYC 370

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS109
General Botany

Prerequisites: BIOS 101 and 101L or equivalent.

Description: Introduction to the plant kingdom and to plants as biological organisms; structure and function of cells, tissues, and organs with emphasis on seed plants; the important processes and concepts of classification, inheritance, evolution, and ecology.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS110
Human Biology

Prerequisites: Parallel registration in BIOS 110L.

High school chemistry or equivalent strongly recommended. Credit toward the degree cannot be earned in both BIOS 101 and BIOS 110. BIOS 110 does not count in the Biological Sciences major.

Description: Introduction to biology with a focus on organization of molecules and cells to the level of human body systems; basic structure (anatomy) and function (physiology) of human tissues, organs and organ systems; reproduction, genetics; DNA technology and genetic engineering.

This course is a prerequisite for: BIOS 110L

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
Offered:FALL/SPR

Credit Hours:3

ACE:

BIOS110L
Human Biology Laboratory

Prerequisites: Parallel registration in BIOS 110.

High school chemistry or equivalent strongly recommended. Credit toward the degree cannot be earned in both BIOS 101L and BIOS 110L. BIOS 110L does not count in the Biological Sciences major.

Description: Hands-on lab exercises to understand biological concepts of human organization from molecules to cells to the body systems, basic structure and function of human tissues, organs, organ systems, reproduction, genetics, as well as DNA technology and genetic engineering.

This course is a prerequisite for: BIOS 110

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB
Offered:FALL/SPR

Credit Hours:1

ACE:

BIOS111
The Biology of Microorganisms

Prerequisites: BIOS 101 and 101L, or equivalent.

Description: Comparative study of microorganisms, principles and applications.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS112
Introduction to Zoology

Prerequisites: Must enroll in both BIOS 112 (lecture) and BIOS 112L (lab)

Description: Survey of the animal kingdom with emphasis on the evolution, ecology, and behavior of major animal groups.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS112L
Introduction to Zoology Lab

Prerequisites: Must enroll in both BIOS 112 (lecture) and BIOS 112L (lab)

Description: Laboratory exercises and experiments that complement material covered in BIOS 112.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS115
Insect BiologyCrosslisted with ENTO 115

Description: Fundamental insect biology (anatomy, development, physiology, behavior, ecology and diversity). Economic and medical importance of insects and principles of insect pest management.

This course is a prerequisite for: ENTO 200; ENTO 400; FORS 411

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 4 Science

Credit Hours:3

ACE:ACE 4 Science

BIOS116
Insect IdentificationCrosslisted with ENTO 116

Description: Identification of representative orders and families of insects by their anatomy, metamorphosis, habits and habitats. Sight recognition emphasized but dichotomous keys also used. Interrelation of insect and habitats stressed.

This course is a prerequisite for: FORS 411

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS117
Life in the UniverseCrosslisted with ASTR 117, GEOL 117

Description: Survey of what modern science tells us about the possibilities of life elsewhere in the universe. Topics include how the Earth formed and became suitable for life, how life arose on the Earth, the conditions under which life can thrive, places in the solar system that might support life, the existence of other solar systems that might provide suitable habitats, and attempts to find evidence of life on other planets.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 4 Science

Credit Hours:3

ACE:ACE 4 Science

BIOS136
Discovery Research: Virus Hunting

Prerequisites: By permission

Description: Perform original research by using the scientific method to isolate a virus that infects a harmless bacterium (bacteriophage) from local soil samples. Lab skills acquired include pipetting, aseptic technique, and serial dilutions; use basic DNA and electron microscopy analyses to characterize the phage.

This course is a prerequisite for: BIOS 137

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LAB
Offered:FALL

Credit Hours:2

ACE:

BIOS137
Discovery Research: Virus Genome Analyses

Prerequisites: BIOS 136 and by permission.

Description: Build on an original project involving isolation of a virus that infects a harmless bacterium (bacteriophage) using bioinformatic tools to analyze and annotate the sequenced bacteriophage genome.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LAB
Offered:SPRING

Credit Hours:2

ACE:

BIOS160
Introduction to Clinical Laboratory Science

Description: Introduces the pre-clinical laboratory scientist/medical technologist to the profession of clinical laboratory science. Includes lessons in ethics, organization of the medical team, professionalism, automation, medical terminology, hematology, blood bank, clinical chemistry, and medical microbiology. For students interested in a career in clinical laboratory science/medical technology. BIOS 160 will not count toward a major in biological sciences.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LEC

Credit Hours:1

ACE:

BIOS189H
University Honors Seminar

Prerequisites: Good standing in the University Honors Program or by invitation.

University Honors Seminar 189H is required of all students in the University Honors Program.

Description: Topic varies.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 4 Science

Credit Hours:3

ACE:ACE 4 Science

BIOS191
Biological Sciences Learning Community Freshman Seminar

Prerequisites: Open to Biological Sciences Learning Community students only

Description: An exploration of biological sciences for undergraduates in the Biological Sciences Learning Community. Topics vary.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LEC

Credit Hours:1

ACE:

BIOS203
Bioethics

Prerequisites: Sophomore standing; BIOS 101 and 101L.

Description: Relevance of biological science to society and its environment examined through readings, guest lecturers, and discussion.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS205
Genetics, Molecular and Cellular Biology Laboratory

Prerequisites: BIOS 206 or parallel

Description: Series of lab exercises to introduce principles of genetic, molecular and cellular biology. Experiments done using model systems to identify, map and clone genes; analyze gene products and expression; and fractionate cell components.

This course is a prerequisite for: BIOS 302; FORS 401

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LEC

Credit Hours:2

ACE:

BIOS206
General Genetics

Prerequisites: LIFE 120 & LIFE 120L and LIFE 121 & LIFE 121L

Description: Inheritance and regulation of genes in organisms and populations. Fundamentals of genomics and bioinformatics.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS207
Ecology and Evolution

Prerequisites: LIFE 120 & LIFE 120L and LIFE 121 & LIFE 121L

Description: Introduction to the principles and processes of ecology and evolution. Structure and dynamics of populations and communities; biotic and abiotic interactions; mechanisms of evolutionary change; natural selection; adaptation; and speciation.

This course is a prerequisite for: BIOS 472

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS213
Human Physiology

Prerequisites: BIOS 101 & BIOS 101L or LIFE 120 & LIFE 120L or equivalent; parallel BIOS 213L

Description: Elementary survey of the basic functional systems of the human body: the muscular, nervous, receptor, circulatory, respiratory, digestive, excretory, endocrine, and reproductive systems.

This course is a prerequisite for: ASCI 341; NUTR 355; VBMS 303; VBMS 403; VBMS 410

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS213L
Human Physiology

Prerequisites: BIOS 101 & BIOS 101L or LIFE 120 & LIFE 120L or equivalent; parallel BIOS 213

Description: Laboratory exercises and experiments that complement material covered in BIOS 213.

This course is a prerequisite for: VBMS 303; VBMS 403

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS214
Human Anatomy

Prerequisites: Sophomore standing.

Cadaver prosections are studied in the lab. BIOS 214 is 'Letter Grade Only'.

Description: Introduction to the major organ systems of the human body including skeletal, major muscle, nervous, digestive, circulatory, excretory, and reproductive systems. Anatomical structures as they pertain to clinical anatomy.

This course is a prerequisite for: NUTR 384

Course details
Credit Hours:5
Max credits per semester:5
Max credits per degree:5
Course Format:LEC

Credit Hours:5

ACE:

BIOS216
Plant Breeding Principles and PracticeCrosslisted with AGRO 216, HORT 216

Prerequisites: High school biology and chemistry. BIOS 101 and 101L, or 102 or equivalent recommended.

Description: Plant breeding theory and technique. Application of genetic principles to plant improvement. Experience with breeding agronomic and horticultural plant species to illustrate plant mating systems and breeding principles.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LEC

Credit Hours:2

ACE:

BIOS220
Principles of EcologyCrosslisted with NRES 220

Prerequisites: 4 hrs biological sciences, and MATH 101 or 103.

BIOS 220 is not open to students who have completed BIOS 207. BIOS 220 will not count toward a major in BIOS.

Description: Ecology as a quantitative discipline that integrates the life and earth sciences to understand the dynamics of natural and managed ecosystems.

This course is a prerequisite for: NRES 311; NRES 862, NRES 462

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS222
Ecology LaboratoryCrosslisted with NRES 222

Prerequisites: NRES/BIOS 220 or parallel.

Description: Field and laboratory experiments in terrestrial and aquatic ecology.

This course is a prerequisite for: NRES 862, NRES 462; WATS 481, WATS 881, BIOS 481, NRES 481

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS232
Ecological Issues in the Great Plains

BIOS 232 is not open to students who have completed BIOS 22. BIOS 232 will not count toward a major in biological sciences.

Description: Basic concepts in ecology, including comparison of major world ecosystems, especially the Great Plains. Interplay of ecological principles and human activities.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 8 Civic/Ethics/Stewardship

Credit Hours:3

ACE:ACE 8 Civic/Ethics/Stewardship

BIOS237
Basic Applications of Bioinformatics

Prerequisites: LIFE 121 and LIFE 121L

Description: Provides a broad overview of bioinformatics. Shows how bioinformatics can help solving problems in biological research. Covered topics: biological databases, molecular biology tools, sequence comparison methods, phylogenetic inference, and molecular graphics.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS280
The Pre-Health Experience for Biological Sciences Majors

Prerequisites: Sophomore standing; BIOS major; BIOS 213 or 214.

Description: Exposure for pre-medical and other pre-health science students to career in health sciences. Shadow a professional and research medically-based careers.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LEC

Credit Hours:1

ACE:

BIOS295
Topics in Biology

Prerequisites: Permission.

Description: Topic varies.

Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:1-3

ACE:

BIOS296
Independent Study in Biology

Prerequisites: 4 hrs BIOS and permission.

A maximum of 3 credit hours may be counted toward the major in BIOS. Before registering, arrangements must be made with a faculty member in BIOS to reach an agreement on the scope and determine the amount of credit for the project.

Description: Opportunity to participate in work in a research laboratory in order to gain some insight into the philosophy and methods of original research.

Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Course Format:IND

Credit Hours:1-3

ACE:

BIOS300
Toxins in the EnvironmentCrosslisted with ENTO 300, NRES 300

Prerequisites: One semester BIOS and one semester CHEM

Description: Introduction to the principles of toxicology as they apply to environmental contaminants, agri-chemicals, and industrial and naturally occurring chemicals.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS302
Cell Biology

Prerequisites: BIOS 206; one semester organic chemistry. BIOS 205 or equivalent, and two semesters organic chemistry recommended

Description: The design, execution, and evaluation of scientific experiments that advance the knowledge of cell and molecular biology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS303
Molecular Biology

Prerequisites: BIOS 206

Description: Molecular biology of prokaryotes and eukaryotes. Review of the experimental basis for the principles of the discipline.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS310
School of Biological Sciences Seminar

Prerequisites: LIFE 120 and LIFE 121

P/N only.

Description: Reviews of current literature of general interest; reports of research activities by staff and guest speakers.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:3
Course Format:LEC

Credit Hours:1

ACE:

BIOS312
Microbiology

Prerequisites: LIFE 120 and LIFE 121, one year general chemistry, and one semester organic chemistry or one semester biochemistry. BIOS 206 recommended.

Parallel registration in BIOS 313 or 314 recommended.

Description: Microbial cell structure, genetics, metabolic and biosynthetic activity, diversity, ecology and evolution including host-microbe interactions.

This course is a prerequisite for: BIOS 421, BIOS 821, MBIO 421; BIOS 440, BIOS 840, VBMS 840, MBIO 440; VBMS 403

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS313
Molecular Microbiology Laboratory

Prerequisites: LIFE 120 and LIFE 121, one year general chemistry; one semester organic chemistry or one semester biochemistry BIOS 206 recommended. Parallel registration in BIOS 312 recommended.

Credit towards the degree may not be earned in both BIOS 313 and 314.

Description: Microbiology techniques which include recombinant DNA methods used in industry, medicine and research.

This course is a prerequisite for: BIOS 440, BIOS 840, VBMS 840, MBIO 440

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LAB

Credit Hours:2

ACE:

BIOS314
Microbiology Laboratory

Prerequisites: LIFE 120 and LIFE 121, one year general chemistry; one semester organic chemistry or one semester biochemistry BIOS 206 recommended.

Parallel registration in BIOS 312 recommended. Credit towards the degree may not be earned in both BIOS 313 and 314.

Description: Traditional microbiology techniques without recombinant DNA methods.

This course is a prerequisite for: BIOS 440, BIOS 840, VBMS 840, MBIO 440

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS315
Vertebrate Embryology

Prerequisites: BIOS 206

Description: Gametogenesis, fertilization, cleavage, early development of a number of vertebrates, and the development of specific organ systems. Includes a three-hour lab in which the morphological aspects of development are illustrated on slides and in which some modern techniques used in experimental mammalian development are introduced.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS316
Case Studies in Theoretical EcologyCrosslisted with MATH 316, NRES 316

Prerequisites: Permission.

Case studies are structured around preparation for subsequent independent research (BIOS 498 or MATH 496).

Description: Introduction to biological literature, applied mathematics, computer programming, and/or statistical techniques relevant to particular questions in ecology, evolution, and behavior. Typical mathematical topics include discrete dynamics, systems of differential equations, matrix algebra, or statistical inference and probability.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
Groups:Advanced Mathematics Courses

Credit Hours:3

ACE:

BIOS317
The Biology of Plants

Prerequisites: LIFE 120 and LIFE 121

Field trips are required and may occur outside of scheduled class time.

Description: Introduction to the basic principles and concepts of the biology of plants. Adaptive variation and biodiversity of plants considering the relationships of plant structure to function integrating across succeeding levels of organization: molecule, cell, tissue, organism, organism, population, community, and ecosystem.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS326
Biology of Viruses

Prerequisites: BIOS 206; one year general chemistry and one semester organic chemistry

Description: Fundamental concepts in virology including basic features of structure, evolution, diseases, replication cycles and virus-host interactions.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS369
Introductory Plant PathologyCrosslisted with PLPT 369

Prerequisites: BIOS 101 and 101L, or 109.

Description: Introduction to the study of plant diseases including relation of plant disease to crop production, environment, and man. Examples and demonstrations emphasize horticultural and agronomic crops of Nebraska.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS381
Invertebrate Zoology

Prerequisites: LIFE 121 & LIFE 121L

Description: Comparative study of the morphology and natural history of invertebrate animals; emphasis on phylogenetic relationships.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS385
Parasitology

Prerequisites: LIFE 121 & LIFE 121L

Description: Emphasis on parasitic diseases of humans. Impact of parasitism on societies considered in addition to the clinical consequences for infected individuals. Means of transmission, diagnosis, and treatment considered in respect to recent technological advances in production of monoclonal antibodies and genetic engineering. Nature and biological significance of parasitism are viewed in terms of prospects for control.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS386
Vertebrate ZoologyCrosslisted with NRES 386

Prerequisites: LIFE 121 & LIFE 121L

Description: Evolutionary origin and relationships, natural history, and ecological adaptations of vertebrates. Comparative form and function, particularly of bone and muscle systems among and the diversity within vertebrate groups.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS388
Comparative Anatomy of the Vertebrates

Prerequisites: LIFE 121 & LIFE 121L

Description: Evolutionary development and comparative structure of the chordate organ systems, including dissections of the dogfish, salamander, and cat.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS389
Anatomy and/or Physiology Internship

Description: A combination of academic work and instruction in the anatomy or physiology laboratories in biological sciences: cadaver dissection or work with physiological equipment; assist in the instruction of anatomical and physiological concepts. Open only to students who expect to become teaching assistants in anatomy or physiology

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LEC

Credit Hours:1

ACE:

BIOS394
Seminar in Behavioral BiologyCrosslisted with PSYC 394

Prerequisites: PSYC 273 OR PSYC 373/BIOS 373

Description: Critical reading and discussion of literature on topics dealing with the biological bases of behavior.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:24
Course Format:LEC

Credit Hours:1

ACE:

BIOS395
Internship

Description: Combination of work outside the University and academic work in biological sciences arranged through the Career Services Office. Specifics of requirements to be arranged with supervising faculty member. BIOS 395 will not count toward a major in BIOS.

Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Course Format:IND

Credit Hours:1-3

ACE:

BIOS398H
Honors Seminar

Prerequisites: Enrollment in the biological sciences honors program

Description: Special topics in biology.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LEC

Credit Hours:1

ACE:

BIOS399H
Honors Research

Prerequisites: Open to candidates for degrees with distinction or enrollment in the biological sciences honors program.

Description: Independent research leading to an honors thesis and exam in accordance with the College's degrees with distinction procedure.

Course details
Credit Hours:1-4
Max credits per semester:4
Max credits per degree:4
Course Format:IND

Credit Hours:1-4

ACE:

BIOS402
Cancer BiologyCrosslisted with BIOS 802

Prerequisites: BIOS 206 and Senior standing

Description: Principles of cancer genetics, cancer prevention, and new methods for diagnosis and therapy. Fundamentals of the cell and molecular events that lead to human cancer.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS406
Insect EcologyCrosslisted with BIOS 806, ENTO 406, ENTO 806

Prerequisites: BIOS/NRES 220 and 222.

Description: Biotic and abiotic factors as they influence insect development, behavior, distribution, and abundance.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS407
Biology of Cells and OrganellesCrosslisted with BIOS 807

Prerequisites: BIOS 206

Description: Regulation and timing of macromolecular synthesis during the cell cycle; the genetic autonomy of mitochondria and chloroplasts.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS408
Functional HistologyCrosslisted with BIOS 808, VBMS 408, VBMS 808

Prerequisites: BIOS 101 and 101L or LIFE 120 and 120L or BIOS 112; BIOS 213 or ASCI 240. BIOS 315 recommended.

Description: Microscopic anatomy of the tissues and organs of major vertebrate species, including humans. Normal cellular arrangements of tissues and organs as related to their macroscopic anatomy and function, with reference to sub-cellular characteristics and biochemical processes. Functional relationships among cells, tissues, organs and organ systems, contributory to organismal well being. General introduction to pathological processes and principles underlying some diseases.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS412
Human GeneticsCrosslisted with BIOS 812

Prerequisites: BIOS 206 and Senior standing

Description: Genetic basis of human variation, with emphasis on methods of applying genetic principles to humankind. Genetic ratios in pooled data; population and quantitative genetics; consanguinity; polygenic inheritance; blood types; sex linkage; linkage and crossing over; sex determination; visible chromosome variation; mutation; heredity and environment; eugenics; anthropological genetics; molecular genetics and molecular basis of disease; human genome project.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS415
Developmental BiologyCrosslisted with BIOS 815

Prerequisites: BIOS 206

Description: Survey of topics in developmental biology, both animal and plant development.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS416
Biodiversity ConservationCrosslisted with BIOS 816

Description: Basic conservation science theory and conservation decision making tools which are essential for making effective decisions for biodiversity conservation. Topics include systematic conservation planning, population viability analysis, risk assessment, and applying those tools to real conservation problems.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS418
Advanced GeneticsCrosslisted with BIOS 818

Prerequisites: BIOS 206 and Senior standing

Description: In-depth study of the principles and methodology of genetics, with emphasis on Drosophila: multiple alleles and complex loci, linkage and recombination, chromosome rearrangements, fine structure analysis, sex determination, recombinant DNA, and gene function in development.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS419
Behavioral NeuroscienceCrosslisted with BIOS 819, PSYC 465, PSYC 865

Prerequisites: PSYC 273 or PSYC 373 or BIOS 373.

Description: Relationship of physiological variables to behavior, an introduction to laboratory techniques in neuropsychology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS420
Molecular GeneticsCrosslisted with BIOS 820, VBMS 820, MBIO 420

Prerequisites: BIOS 206 and Senior standing

Description: Molecular basis of genetics. Gene structure and regulation, transposable elements, chromosome structure, DNA replication, and repair mechanisms and recombination.

This course is a prerequisite for: FDST 908B

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS421
Microbial DiversityCrosslisted with BIOS 821, MBIO 421

Prerequisites: BIOS 206 and BIOS 312 and Senior Standing.

Description: Diversity of microbial cell composition, structure, and function enabling movement, metabolism, symbiosis, and adaptation using bacterial, fungal, algal, and viral examples. A physiological, biochemical and molecular approach used throughout.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS422
Comparative PhysiologyCrosslisted with BIOS 822

Prerequisites: BIOS 213

Description: Comprehensive survey of comparative physiology with emphasis on the diversity of adaptations in basic physiological systems and the effects of environmental parameters upon such systems. Comparative physiology of osmoregulation, temperature regulation, metabolism, muscle, central nervous function, and sensory function.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS422L
Comparative Physiology Laboratory

Prerequisites: Co-enrollment with BIOS 422

Letter grade only.

Description: Physiological adaptations in ecological and evolutionary context.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS425
Plant BiotechnologyCrosslisted with BIOS 825

Prerequisites: BIOS 206

Description: Introduction to the use of plants for basic and applied purposes by deliberate manipulation of their genomes; techniques in plant genetic engineering; manipulations of plant development and metabolism; engineering pest, disease, and stress resistance; plants as bioreactors; and environmental and social impacts of plant biotechnology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS426
Systems BiologyCrosslisted with BIOS 826

Prerequisites: LIFE 120 and LIFE 121, or BIOS 101, or equivalent; STAT 218 or equivalent; BIOS 206 and CSCE 155T are recommended, but not required.

Description: Fundamentals of the analysis of high throughput experiments to understand complex biological systems. Principles and methods such as next generation sequencing, protein-protein interaction networks, regulatory networks, and biological data mining and integration. Emerging research in new biotechnology and data analysis in biomedical and life sciences.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS427
Practical Bioinformatics LaboratoryCrosslisted with BIOS 827

Prerequisites: BIOS 206 or equivalent.

Description: Basic knowledge and skills needed for general bioinformatics, genomics and proteomics analyses. Various computational analyses including database search, sequence alignment, phylogenetic reconstruction, gene prediction/mining, microarray data analyses and protein structure analyses.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS428
Perl Programming for Biological ApplicationsCrosslisted with BIOS 828

Prerequisites: LIFE 120 and LIFE 121

Description: Computer programming, using Perl, as applied to biological sciences, bioinformatics, computational biology, and genomics.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS429
Phylogenetic BiologyCrosslisted with BIOS 829

Prerequisites: BIOS 207 and Senior standing

Description: Principles of phylogenetic inference and emphasis on the application of phylogenetic hypotheses in biology and the biomedical sciences. How inferences derived from phylogenetic trees can be applied in different areas of biological investigation including systematics, biogeography, conservation biology, molecular evolution, genome structure, epidemiology, population biology, ecology, character evolution, behavior, and macroevolution.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS431
Structure and MetabolismCrosslisted with BIOC 431, BIOC 831, BIOS 831, CHEM 431, CHEM 831

Prerequisites: CHEM 252 or CHEM 262 with a grade of C or better. LIFE 120 and BIOS 206 are recommended

First course of a two-semester, comprehensive biochemistry course sequence.

Description: Structure and function of proteins, nucleic acids, carbohydrates and lipids; nature of enzymes; major metabolic pathways of catabolism; and biochemical energy production.

This course is a prerequisite for: VBMS 410

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS432
Metabolism and Biological InformationCrosslisted with BIOC 432, BIOC 832, CHEM 432, CHEM 832, BIOS 832

Prerequisites: BIOC 431/831 with a grade of C or better, BIOS 206 or AGRO 215.

Continuation of BIOC 431/831.

Description: Major metabolic pathways of anabolism, structural and biochemical aspects of biological information flow and use in biotechnology.

This course is a prerequisite for: BIOC 435; BIOC 934, BIOS 934, CHEM 934

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS433
Biochemistry LaboratoryCrosslisted with BIOC 433, BIOC 833, BIOS 833, CHEM 433, CHEM 833

Prerequisites: BIOC 431/831 (or concurrent enrollment) or CHEM 435/835.

Description: Introduction to techniques used in biochemical and biotechnology research, including measurement of pH, spectroscopy, analysis of enzymes, chromatography, fractionation of macromolecules, electrophoresis, and centrifugation.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LEC

Credit Hours:2

ACE:

BIOS434
Plant BiochemistryCrosslisted with AGRO 434, BIOC 434, CHEM 434, AGRO 834, BIOC 834, BIOS 834, CHEM 834

Prerequisites: BIOC/BIOS/CHEM 431/831.

Description: Biochemical metabolism unique to plants. Relationships of topics previously acquired in general biochemistry to biochemical processes unique to plants. Biochemical mechanisms behind physiological processes discussed in plant or crop physiology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS436
Quaternary Paleoclimatology and PaleoecologyCrosslisted with BIOS 836, GEOL 423, GEOL 823

Prerequisites: 12 hrs GEOL or BIOS.

Description: Analysis and interpretation of the Quaternary period's paleoecological data. Patterns of long-term climate variation. Distribution patterns and responses of organisms and ecosystems to Quaternary environmental change.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS437
Research Techniques in BiochemistryCrosslisted with BIOC 437, BIOC 837, BIOS 837

Prerequisites: BIOC/BIOS/CHEM 433/833, or permission

Description: Methods approach to systems biology analysis. Molecular identification and quantification employing techniques such as mass spectrometry, chromatography, electrophoretic fractionation, transcriptomics, protemics and metabolomics. Data and pathway analysis with computational methods.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS438
Biogeochemical CyclesCrosslisted with BIOS 838, GEOL 424, GEOL 824

Prerequisites: CHEM 109 or 113; 12 hrs geology or biological sciences.

Description: Chemical cycling at or near the earth's surface, emphasizing interactions among the atmosphere, biosphere, geosphere and hydrosphere. Modern processes, the geological record, and human impacts on elemental cycles.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS439
Dynamics of Biochemical and Biological NetworksCrosslisted with BIOC 439, BIOC 839, BIOS 839

Prerequisites: BIOS 206, BIOC 321 or BIOC 431 (or equivalent)

Description: To introduce and integrate, students in biochemistry and other life sciences, to the field of computational modeling of biochemical and biological network systems into a seamless curriculum.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS440
Microbial PhysiologyCrosslisted with BIOS 840, VBMS 840, MBIO 440

Prerequisites: BIOS 312 and either 313 or 314, or permission

Description: Molecular approaches to the study of prokaryotic cell structure and physiology, including growth, cell division, metabolism, and alternative microbial life styles.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS441
Pathogenic MicrobiologyCrosslisted with BIOS 841, VBMS 441, VBMS 441H, VBMS 841

Prerequisites: BIOS 312 and either 313 or 314, or permission.

Description: Fundamental principles involved in host-microorganism interrelationships. Identification of pathogens, isolation, propagation, mode of transmission, pathogenicity, symptoms, treatment, prevention of disease, epidemiology, and methods of control.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS442
EndocrinologyCrosslisted with ASCI 442, ASCI 842, BIOS 842, VBMS 842

Prerequisites: A course in vertebrate physiology and/or biochemistry.

Description: Mammalian endocrine glands from the standpoint of their structure, their physiological function in relation to the organism, the chemical nature and mechanisms of action of their secretory products, and the nature of anomalies manifested with their dysfunction.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS443
ImmunologyCrosslisted with BIOS 843, VBMS 843, MBIO 443

Prerequisites: BIOS 206 and one semester organic chemistry

Description: Fundamental consideration of cellular and humoral mechanisms of immunity, the structure and function of immunoglobulins, antigen-antibody interactions; hypersensitivity; transplantation and tumor immunity; immune and autoimmune disorders.

This course is a prerequisite for: VBMS 910

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS444
GeomicrobiologyCrosslisted with BIOS 844, GEOL 444, GEOL 844

Prerequisites: 3 hours biological sciences and 3 hours chemistry.

Description: Lectures and discussions of primary literature regarding microorganisms and their role transforming Earth through geologic time.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS445
Food MicrobiologyCrosslisted with BIOS 845, FDST 405, FDST 805

Prerequisites: BIOS 312; CHEM 251; BIOC 321.

Description: Nature, physiology, and interactions of microorganisms in foods. Introduction to food-borne diseases, the effect of food processing systems on the microflora of foods, principles of food preservation, food spoilage, and foods produced by microorganisms. Food plant sanitation and criteria for establishing microbial standards for food products.

This course is a prerequisite for: FDST 460, FDST 860; FDST 908B

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS446
Food Microbiology LaboratoryCrosslisted with BIOS 846, FDST 406, FDST 806

Prerequisites: Parallel in FDST 405/805/BIOS 446/846.

Description: The microorganisms in foods and the methods used to study them.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LAB

Credit Hours:2

ACE:

BIOS447
Soil MicrobiologyCrosslisted with AGRO 460, NRES 460, SOIL 460, AGRO 860, BIOS 847, NRES 860

Prerequisites: One semester microbiology; one semester biochemistry or organic chemistry.

Description: Soil from a microbe's perspective-growth, activity and survival strategies; principles governing methods to study microorganisms and biochemical processes in soil; mechanisms controlling organic matter cycling and stabilization with reference to C, N, S, and P; microbial interactions with plants and animals; and agronomic and environmental applications of soil microorganisms.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS448
Human Growth and DevelopmentCrosslisted with ANTH 448, ANTH 848, BIOS 848

Prerequisites: ANTH 242 and 242L, or BIOS 101 and 101L.

Description: Biological diversity from an evolutionary perspective. The history of the study of human physical growth and biological principles of growth. Genetic, epigenetic and hormonal effects on human and other mammal growth patterns, and environmental factors that influence growth. Effects of nutrition, disease, socio-economic status, pollution, etc. Unique features of human growth in its various stages. How anthropologists interpret variation in growth patterns among human populations and the possible adaptive significance of this variation.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
Groups:Biological Anthropology

Credit Hours:3

ACE:

BIOS450
Biology of Wildlife PopulationsCrosslisted with BIOS 850, NRES 450, NRES 850

Prerequisites: NRES 311; MATH 104 or above; STAT 218 or equivalent

Description: Principles of population dynamics. Management strategies (for consumptive and nonconsumptive fish and wildlife species) presented utilizing principles developed.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
Offered:SPRING

Credit Hours:4

ACE:

BIOS452
Field EpidemiologyCrosslisted with BIOS 852

Prerequisites: LIFE 120 and LIFE 121 and one additional 200-level life science course; or permission Offered summers only at Cedar Point Biological Station

Description: Principles of epidemiology and the role in modern medicine. Combination of theory and practice with living populations.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS453
Predator EcologyCrosslisted with BIOS 853

Prerequisites: BIOS 207 or NRES 220

Description: Conservation and management of native and invasive predators. Functional and numerical responses. Evolution of predator-prey interactions. Optimal foraging. Modeling predator-prey population dynamics. Trophic cascades. Prey defenses against predation.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS454
Ecological InteractionsCrosslisted with BIOS 854, NRES 454, NRES 854

Prerequisites: LIFE 121 & 121L, BIOS 207 or BIOS 220 and Senior standing

Description: Nature and characteristics of populations and communities. Interactions within and between populations in community structure and dynamics. Direct and indirect interactions and ecological processes, competition, predation, parasitism, herbivory, and pollination. Structure, functioning and persistence of natural communities, foodweb dynamics, succession, and biodiversity.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS455
Great Plains FloraCrosslisted with BIOS 855

Prerequisites: LIFE 120 and LIFE 121 BIOS 207 or BIOS 220 recommended

Description: Plant identification. Field study of the flora in various habitats. Field trips include grassland and woodland vegetation of this region.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS456
Mathematical Models in BiologyCrosslisted with BIOS 856, NRES 456

Prerequisites: Junior standing; major in the biological sciences; MATH 106 or 107.

Description: Biological systems, from molecules to ecosystems, are analyzed using mathematical techniques. Strengths and weaknesses of mathematical approaches to biological questions. Brief review of college level math; introduction to modeling; oscillating systems in biology; randomness in biology; review of historically important and currently popular models in biology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS457
Ecosystem EcologyCrosslisted with BIOS 857, GEOL 457, GEOL 857

Prerequisites: BIOS 207 and CHEM 110 and Senior standing

Description: Processes controlling the cycling of energy and elements in ecosystems and how both plant and animal species influence them. Human-influenced global and local changes that alter these cycles and ecosystem functioning.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS458
WetlandsCrosslisted with NRES 468, NRES 868, WATS 468

Prerequisites: 12 hrs biological sciences; BIOS 220; CHEM 109 and 110.

Description: Physical, chemical and biological processes that occur in wetlands; the hydrology and soils of wetland systems; organisms occurring in wetlands and their ecology wetland creation, delineation, management and ecotoxicology.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS459
LimnologyCrosslisted with BIOS 859, NRES 459, NRES 859, WATS 459

Prerequisites: 12 hrs BIOS, including BIOS/NRES 220/BIOS220x; two semesters CHEM.

Description: Physical, chemical, and biological processes that occur in fresh water. Organisms occurring in fresh water and their ecology; biological productivity of water and its causative factors; eutroplication and its effects.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS462
Animal BehaviorCrosslisted with BIOS 862

Prerequisites: BIOS 206, 207 and Senior Standing

Description: Introduction to animal behavior stressing the ethological approach. Anatomical and physiological bases of behavior, ontogenetic and phylogenetic observations, and the relations of animal behavior studies to genetics, ecology, taxonomy, and evolution.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS464
Fisheries BiologyCrosslisted with BIOS 864, NRES 464, NRES 864

Prerequisites: BIOS/NRES 489/889 or equivalent.

Description: Biology of fishes. Factors that affect fishes in the natural environment. Techniques used in the analysis and management of fish populations.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS468
Field Animal BehaviorCrosslisted with BIOS 868

Prerequisites: LIFE 120 and LIFE 121 BIOS 207 or BIOS 220 recommended

Offered summers only at Cedar Point Biological Station.

Description: Behavior of animals. Stresses methods for testing evolutionary hypotheses under field conditions with emphasis on foraging behavior, animal communication, and animal social systems.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS470
Prairie EcologyCrosslisted with BIOS 870

Prerequisites: BIOS 207 or equivalent.

Description: Structure, function, and distribution of communities. Interaction of different species with their biotic and abiotic environments.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS471
Plant SystematicsCrosslisted with BIOS 871

Prerequisites: LIFE 121 and LIFE 121L

Description: Overview of the diversity of plants and algae, with emphasis on phylogenetic relationships, the evolution of important physical and genomic characteristics, principles of plant classification and identification, and modern methods of plant molecular systematics. Lab work on taxonomic analysis and plant identification.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS472
Evolution

Prerequisites: BIOS 207 and Senior standing

Description: The principles and processes of micro- and macroevolution. Mechanisms behind evolutionary change and examples of these processes in a wide variety of organisms.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:3

ACE:ACE 10 Integrated Product

BIOS474
HerpetologyCrosslisted with BIOS 874, NRES 474, NRES 874

Prerequisites: BIOS/NRES 386 and permission. BIOS 388 recommended.

Description: Fossil and living amphibians and reptiles. Anatomy, classification, ecology and evolution.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS475
Avian BiologyCrosslisted with BIOS 875

Prerequisites: LIFE 121 & LIFE 121L

May also be offered at Cedar Point Biological Station.

Description: Biology of birds emphasizing the behavior and ecology of this group. Topics include avian diversity, systematics & evolutionary history, flight, foraging, migration, communication, reproductive biology, population ecology and conservation biology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS475L
Avian Biology LaboratoryCrosslisted with BIOS 875L

Prerequisites: Parallel BIOS 475/875 and permission

An immersive field biology experience at Cedar Point Biological Station.

Description: Avian field identification in diverse prairie, riparian, and montane habitats. Individual studies of foraging behavior, territoriality, anti-predator behavior, mating systems, or nesting ecology.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS476
MammalogyCrosslisted with BIOS 876, NRES 476, NRES 876

Prerequisites: 8 hrs BIOS; BIOS/NRES 386 or NRES 311.

May also be offered at Cedar Point Biological Station. Field trips are required and may occur outside of scheduled class time. Lab and field time emphasize diversity of mammalian families and species identification of Nebraska mammals.

Description: Evolution, natural history, ecology, and functional morphology of planetary mammals and mammals of the Northern Great Plains.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS477
Bioinformatics and Molecular EvolutionCrosslisted with BIOS 877

Prerequisites: BIOS 206 or parallel and CHEM 251 or equivalent

Description: Pairwise and multiple alignments, sequence similarity and domain search, distance estimation, phylogenetic methods, gene mining, protein classification and structure. Algorithms used in bioinformatics as well as fundamental concepts of molecular evolution that underlie various bioinformatics methods.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS478
Plant AnatomyCrosslisted with BIOS 878

Prerequisites: 8 hrs biological sciences, BIOS 109 recommended.

Description: Development, structure, and function of tissues and organs of the higher plants. Relationships of structure to physiology and ecology of plants.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS480
Ecology and Evolution of ArachnidsCrosslisted with BIOS 880

Prerequisites: BIOS 207 or BIOS 220

Description: Ecology and evolutionary biology of living arachnids.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS481
Stream and River EcologyCrosslisted with WATS 481, WATS 881, NRES 481

Prerequisites: NRES 222 or equivalent

Description: Fundamental physical drivers operating in stream and river ecosystems and how those vary in space and time. Major classes of organisms associated with stream ecosystems and their functional roles. Fundamental controls on biotic diversity in stream and river ecosystems and its variance. Major aspects of stream ecosystem function including energy flow and nutrient cycling. Ecosystem services provided by stream and river ecosystems and causes and consequences of human impacts on streams and rivers. Underlying principles of bioassessment and current methods of stream restoration.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS482
Field EntomologyCrosslisted with BIOS 882, ENTO 411, ENTO 811

Prerequisites: 12 hrs biological sciences.

Description: Field course in insect taxonomy and biology emphasizing field collection, specimen preparation, classification, and insect natural history.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS484
Physiology of ExerciseCrosslisted with BIOS 884, NUTR 484, NUTR 884

Prerequisites: 12 hrs biological sciences, including BIOS 213 or equivalent; BIOS 214 or equivalent

Description: Effects of physical activity on the circulatory, respiratory, and other physiological processes.

This course is a prerequisite for: NUTR 486, NUTR 886

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS485
Aquatic InsectsCrosslisted with BIOS 885, ENTO 402, ENTO 802, NRES 402, NRES 802

Prerequisites: 12 hrs biological sciences.

Description: Biology and ecology of aquatic insects.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Course Format:LEC

Credit Hours:2

ACE:

BIOS485L
Identification of Aquatic InsectsCrosslisted with BIOS 885L, ENTO 402L, ENTO 802L, NRES 402L, NRES 802L

Prerequisites: Parallel ENTO 802, NRES 402/802, BIOS 485/885.

Description: Identification of aquatic insects to the family level.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Course Format:LAB

Credit Hours:1

ACE:

BIOS486
Advanced Topics in Biophysical ChemistryCrosslisted with BIOC 486, BIOC 886, BIOS 886, CHEM 486, CHEM 886

Prerequisites: CHEM 471/871 or 481/881.

Description: Applications of thermodynamics to biochemical phenomena, optical properties of proteins and polynucleotides, and kinetics of rapid reactions.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Course Format:LEC

Credit Hours:3

ACE:

BIOS487
Field ParasitologyCrosslisted with BIOS 887

Prerequisites: LIFE 120 and LIFE 121 BIOS 207 or BIOS 220 recommended

Description: Animal host-parasite relationships, epizootiology, ecology, host distribution, classification, and life cycle stages of animal parasites.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC
ACE Outcomes: ACE 10 Integrated Product

Credit Hours:4

ACE:ACE 10 Integrated Product

BIOS488
Natural History of the InvertebratesCrosslisted with BIOS 888

Prerequisites: LIFE 120 and LIFE 121 BIOS 207 or BIOS 220 recommended

Description: Field course in invertebrate community relations stressing on-site observation of community components, natural history, and interactions.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS489
IchthyologyCrosslisted with BIOS 889, NRES 489, NRES 889

Prerequisites: LIFE 120 and LIFE 121

Description: Fishes, their taxonomy, physiology, behavior, and ecology. Dynamics of fish stocks and factors regulating their production.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Course Format:LEC

Credit Hours:4

ACE:

BIOS497
Special Topics in Biological SciencesCrosslisted with BIOS 897

Prerequisites: LIFE 120 and LIFE 121; and permission

Description: Topics vary by term.

Course details
Credit Hours:1-4
Max credits per semester:4
Max credits per degree:24
Course Format:LEC

Credit Hours:1-4

ACE:

BIOS498
Independent Research in Biological SciencesCrosslisted with BIOS 898

Prerequisites: LIFE 120 and LIFE 121; and permission

Four credit hours may be counted toward the undergraduate BIOS major. Before registering, arrangements must be made with a School of Biological Sciences faculty member to reach an agreement on the scope and to determine the amount of credit for the project.

Description: Independent study and laboratory or field investigation of a specific problem.

Course details
Credit Hours:1-8
Max credits per semester:8
Max credits per degree:8
Course Format:IND

Credit Hours:1-8

ACE: