Biological Sciences (BIOS)

BIOS802
Cancer BiologyCrosslisted with BIOS 402

Prerequisites: BIOS 206 and Senior standing

Description: Principles of cancer genetics, cancer prevention, and new methods for diagnosis and therapy. Fundamentals of the cell and molecular events that lead to human cancer.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS803
Principles of Evolution

Description: Micro- and macroevolutionary patterns and processes. Population genetics, evolutionary ecology, speciation, phylogenetic systematics, and biogeographic patterns of extant and extinct taxa.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS804
Principles of Behavioral Ecology

Description: Introduction to the ecology and evolution of animal behavior.

This course is a prerequisite for: BIOS 962

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS805
Principles of Ecology

Description: Ecological principles at the populations, community, and ecosystem levels. Population growth, meta-population dynamics, competitive and predatory interactions, temporal and spatial variation in community food webs, tropic cascades, patterns and mechanisms underlying species diversity, ecosystem processes, nutrient cycling, and global change.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS806
Insect EcologyCrosslisted with BIOS 406, ENTO 406, ENTO 806

Prerequisites: BIOS/NRES 220 and 222.

Description: Biotic and abiotic factors as they influence insect development, behavior, distribution, and abundance.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS808
Functional HistologyCrosslisted with BIOS 408, VBMS 408, VBMS 808

Prerequisites: BIOS 101 and 101L or LIFE 120 and 120L; BIOS 213 or ASCI 240 or ASCI 340.

Description: Microscopic anatomy of the tissues and organs of major vertebrate species, including humans. Normal cellular arrangements of tissues and organs as related to their macroscopic anatomy and function, with reference to sub-cellular characteristics and biochemical processes. Functional relationships among cells, tissues, organs and organ systems, contributory to organismal well being. General introduction to pathological processes and principles underlying some diseases.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$15

Credit Hours:4

ACE:

BIOS809
Professionalism

Pass/No Pass only.

Description: Discussion of skills needed to be a professional scientist including: writing, submitting, editing, and revision of journal articles and grant proposals; preparation or oral and poster presentations; and ethical issues in research and teaching.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Grading Option:Pass No-Pass

Credit Hours:1

ACE:

BIOS812
Human GeneticsCrosslisted with BIOS 412

Prerequisites: BIOS 206 and Senior standing

Description: Genetic basis of human variation, with emphasis on methods of applying genetic principles to humankind. Genetic ratios in pooled data; population and quantitative genetics; consanguinity; polygenic inheritance; blood types; sex linkage; linkage and crossing over; sex determination; visible chromosome variation; mutation; heredity and environment; eugenics; anthropological genetics; molecular genetics and molecular basis of disease; human genome project.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS816
Biodiversity ConservationCrosslisted with BIOS 416

Prerequisites: BIOS 207 or NRES 220

Description: Basic conservation science theory and conservation decision making tools which are essential for making effective decisions for biodiversity conservation. Topics include systematic conservation planning, population viability analysis, risk assessment, and applying those tools to real conservation problems.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$10

Credit Hours:3

ACE:

BIOS818
Advanced GeneticsCrosslisted with BIOS 418

Prerequisites: BIOS 206 and Senior standing

Description: In-depth study of the principles and methodology of genetics, with emphasis on Drosophila: multiple alleles and complex loci, linkage and recombination, chromosome rearrangements, fine structure analysis, sex determination, recombinant DNA, and gene function in development.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS820
Molecular GeneticsCrosslisted with BIOS 420, VBMS 820, MBIO 420

Prerequisites: BIOS 206 and Senior standing

Description: Molecular basis of genetics. Gene structure and regulation, transposable elements, chromosome structure, DNA replication, and repair mechanisms and recombination.

This course is a prerequisite for: AGRO 963, HORT 963, PLPT 963; BIOS 945; BIOS 964, VBMS 964; FDST 908B

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS821
Microbial DiversityCrosslisted with BIOS 421, MBIO 421

Prerequisites: BIOS 206 and BIOS 312 and Senior Standing.

Description: Diversity of microbial cell composition, structure, and function enabling movement, metabolism, symbiosis, and adaptation using bacterial, fungal, algal, and viral examples. A physiological, biochemical and molecular approach used throughout.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS822
Comparative PhysiologyCrosslisted with BIOS 422

Prerequisites: BIOS 213

Description: Comprehensive survey of comparative physiology with emphasis on the diversity of adaptations in basic physiological systems and the effects of environmental parameters upon such systems. Comparative physiology of osmoregulation, temperature regulation, metabolism, muscle, central nervous function, and sensory function.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$10

Credit Hours:3

ACE:

BIOS823
Quaternary Paleoclimatology and PaleoecologyCrosslisted with BIOS 423, GEOL 423, GEOL 823

Prerequisites: 12 hrs GEOL or BIOS.

Description: Analysis and interpretation of the Quaternary period's paleoecological data. Patterns of long-term climate variation. Distribution patterns and responses of organisms and ecosystems to Quaternary environmental change.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS824
Biogeochemical CyclesCrosslisted with BIOS 424, GEOL 424, GEOL 824

Prerequisites: CHEM 109A and 109L or CHEM 113A and 113L; 12 hrs GEOL or BIOS.

Description: Chemical cycling at or near the earth's surface, emphasizing interactions among the atmosphere, biosphere, geosphere and hydrosphere. Modern processes, the geological record, and human impacts on elemental cycles.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS825
Plant BiotechnologyCrosslisted with BIOS 425

Prerequisites: BIOS 206.

Description: Introduction to the use of plants for basic and applied purposes by deliberate manipulation of their genomes; techniques in plant genetic engineering; manipulations of plant development and metabolism; engineering pest, disease, and stress resistance; plants as bioreactors; and environmental and social impacts of plant biotechnology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS826
Systems BiologyCrosslisted with BIOS 426

BIOS 206 and CSCE 155T are recommended, but not required.

Description: Fundamentals of the analysis of high throughput experiments to understand complex biological systems. Principles and methods such as next generation sequencing, protein-protein interaction networks, regulatory networks, and biological data mining and integration. Emerging research in new biotechnology and data analysis in biomedical and life sciences.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS827
Practical Bioinformatics LaboratoryCrosslisted with BIOS 427

Prerequisites: BIOS 206

No computer programming skill is required.

Description: Basic knowledge and skills needed for general bioinformatics, genomics and proteomics analyses. Various computational analyses including database search, sequence alignment, phylogenetic reconstruction, gene prediction/mining, microarray data analyses and protein structure analyses.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS829
Phylogenetic BiologyCrosslisted with BIOS 429

Prerequisites: BIOS 207 and Senior standing

Description: Principles of phylogenetic inference and emphasis on the application of phylogenetic hypotheses in biology and the biomedical sciences. How inferences derived from phylogenetic trees can be applied in different areas of biological investigation including systematics, biogeography, conservation biology, molecular evolution, genome structure, epidemiology, population biology, ecology, character evolution, behavior, and macroevolution.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option

Credit Hours:4

ACE:

BIOS830
Communicating Science through OutreachCrosslisted with BIOS 430

Prerequisites: BIOS 207.

Students must have at least one afternoon available for running a middle school science club (typically between 3-5pm). Background checks required.

Description: Introduction to science communication, formal versus informal science education, and best practices in informal science education. Review of state and national science standards and how students learn. Introduction to informal science practitioners and facilities in Nebraska. Role playing and development and implementation of hands on, inquiry-based science activities. Training in evaluation and assessment.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:FALL/SPR

Credit Hours:3

ACE:

BIOS831
Biochemistry I: Structure and MetabolismCrosslisted with BIOC 431, BIOC 831, BIOS 431, CHEM 431, CHEM 831

Prerequisites: LIFE 120 with a grade of C or better; CHEM 252 or CHEM 262 with a grade of C or better.

BIOS 206 or PLAS 215 is recommended. First course of a two-semester, comprehensive biochemistry course sequence.

Description: Structure and function of proteins, nucleic acids, carbohydrates and lipids; nature of enzymes; major metabolic pathways of catabolism; and biochemical energy production.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:FALL/SPR

Credit Hours:3

ACE:

BIOS832
Biochemistry II: Metabolism and Biological InformationCrosslisted with BIOC 432, BIOC 832, BIOS 432, CHEM 432, CHEM 832

Prerequisites: BIOC 431/831 with a grade of C or better; BIOS 206 or PLAS 215 with a grade of C or better.

Continuation of BIOC 431/831.

Description: Major metabolic pathways of anabolism, structural and biochemical aspects of biological information flow and use in biotechnology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:FALL/SPR

Credit Hours:3

ACE:

BIOS833
Biochemistry LaboratoryCrosslisted with BIOC 433, BIOC 833, BIOS 433, CHEM 433, CHEM 833

Prerequisites: BIOC 431/831 or parallel; or CHEM 435/835.

Description: Introduction to techniques used in biochemical and biotechnology research, including measurement of pH, spectroscopy, analysis of enzymes, chromatography, fractionation of macromolecules, electrophoresis, and centrifugation.

This course is a prerequisite for: BIOC 437, BIOC 837, BIOS 437, BIOS 837; BIOC 898

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Grading Option:Grade Pass/No Pass Option
Offered:FALL/SPR
Course and Laboratory Fee:$50

Credit Hours:2

ACE:

BIOS834
Plant BiochemistryCrosslisted with PLAS 434, BIOC 434, BIOS 434, CHEM 434, AGRO 834, BIOC 834, CHEM 834

Prerequisites: BIOC/BIOS/CHEM 431/831.

Description: Biochemical metabolism unique to plants. Relationships of topics previously acquired in general biochemistry to biochemical processes unique to plants. Biochemical mechanisms behind physiological processes discussed in plant or crop physiology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS835
Evolutionary MedicineCrosslisted with BIOS 435

Prerequisites: BIOS 207 and senior standing

Description: Application of evolutionary tools to biomedical questions. Managing the evolution of drug resistance and pathogen virulence. Evolutionary principles of vaccine design. Emerging infectious disease. Human evolutionary history. Life-history trade-offs in human evolution. Parent-offspring and parent-parent conflict. Mismatch hypothesis. Hygiene hypothesis.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:SPRING

Credit Hours:3

ACE:

BIOS836
MacroecologyCrosslisted with BIOS 436

Prerequisites: BIOS 207

Description: Species-area relationships, latitudinal gradients in species richness, abundance diversity relationships, ecological scaling relationships with body size, community assembly, evolutionary dynamics, climate change, and human impacts on the ecology of the Anthropocene.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS837
Research Techniques in BiochemistryCrosslisted with BIOC 437, BIOC 837, BIOS 437

Prerequisites: BIOC/BIOS/CHEM 433/833.

Description: Methods approach to systems biology analysis. Molecular identification and quantification employing techniques such as mass spectrometry, chromatography, electrophoretic fractionation, transcriptomics, protemics and metabolomics. Data and pathway analysis with computational methods.

This course is a prerequisite for: VBMS 919

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$50

Credit Hours:4

ACE:

BIOS839
Dynamics of Biochemical and Biological NetworksCrosslisted with BIOC 439, BIOC 839, BIOS 439

Prerequisites: BIOS 206 or PLAS 215; BIOC 401 or BIOC 431

Letter grade only.

Description: To introduce and integrate, students in biochemistry and other life sciences, to the field of computational modeling of biochemical and biological network systems into a seamless curriculum.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Graded
Offered:SPRING

Credit Hours:3

ACE:

BIOS840
Microbial PhysiologyCrosslisted with BIOS 440, VBMS 840, MBIO 440, VBMS 440

Prerequisites: BIOS 312; BIOS 313 or BIOS 314.

Description: Molecular approaches to the study of prokaryotic cell structure and physiology, including growth, cell division, metabolism, and alternative microbial life styles.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$20

Credit Hours:3

ACE:

BIOS841
Pathogenic MicrobiologyCrosslisted with BIOS 441, VBMS 441, VBMS 841

Prerequisites: BIOS 312

Description: Fundamental principles involved in host-microorganism interrelationships. Identification of pathogens, isolation, propagation, mode of transmission, pathogenicity, symptoms, treatment, prevention of disease, epidemiology, and methods of control.

This course is a prerequisite for: VBMS 805; VBMS 949

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:SPRING
Course and Laboratory Fee:$25

Credit Hours:3

ACE:

BIOS842
EndocrinologyCrosslisted with ASCI 442, ASCI 842, BIOS 442, VBMS 842

Prerequisites: A course in vertebrate physiology and/or biochemistry.

Description: Mammalian endocrine glands from the standpoint of their structure, their physiological function in relation to the organism, the chemical nature and mechanisms of action of their secretory products, and the nature of anomalies manifested with their dysfunction.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS843
ImmunologyCrosslisted with BIOS 443, VBMS 843, MBIO 443, VBMS 443

Prerequisites: BIOS 206; CHEM 251 or CHEM 255 or CHEM 261.

Description: Fundamental consideration of cellular and humoral mechanisms of immunity, the structure and function of immunoglobulins, antigen-antibody interactions; hypersensitivity; transplantation and tumor immunity; immune and autoimmune disorders.

This course is a prerequisite for: VBMS 852; VBMS 908; VBMS 910; VBMS 948; VBMS 949

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS844
Earth and Environmental MicrobiologyCrosslisted with BIOS 444, GEOL 444, GEOL 844

Prerequisites: 3 hours of BIOS or 3 hours of LIFE; 3 hours of CHEM

Description: An introduction into the role that microorganisms play and have played in natural and man-made environments. Topics covered include microbial diversity and physiology in soil, sediment, and water; microbes in Earth history; biogeochemical cycling; mineral formation and dissolution; biodegradation and bioremediation; biotechnology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS845
Food MicrobiologyCrosslisted with BIOS 445, FDST 405, FDST 805

Prerequisites: BIOS 312

BIOC 401 or BIOC 431 recommended

Description: Nature, physiology, and interactions of microorganisms in foods. Introduction to food-borne diseases, the effect of food processing systems on the microflora of foods, principles of food preservation, food spoilage, and foods produced by microorganisms. Food plant sanitation and criteria for establishing microbial standards for food products.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:FALL/SPR

Credit Hours:3

ACE:

BIOS846
Food Microbiology LaboratoryCrosslisted with BIOS 446, FDST 406, FDST 806

Prerequisites: Parallel in FDST 405/805/BIOS 446/846.

Description: The microorganisms in foods and the methods used to study them.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$40

Credit Hours:2

ACE:

BIOS849
Woody Plant Growth and DevelopmentCrosslisted with HORT 849, NRES 849

Prerequisites: CHEM 251 and AGRO 325

Description: Plant growth and development specifically of woody plants as viewed from an applied whole-plant physiological level. Plant growth regulators, structure and secondary growth characteristics of woody plants, juvenility, senescence, abscission and dormancy.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS850
Biology of Wildlife PopulationsCrosslisted with BIOS 450, NRES 450, NRES 850

Prerequisites: NRES 311; MATH 104 or above; STAT 218 or equivalent

Description: Principles of population dynamics. Management strategies (for consumptive and nonconsumptive fish and wildlife species) presented utilizing principles developed.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Offered:SPRING

Credit Hours:4

ACE:

BIOS851
Invertebrate PaleobiologyCrosslisted with GEOL 451, BIOS 451, GEOL 851

Prerequisites: At least one of: GEOL 103, GEOL 105, LIFE 121

Description: Overview of the key traits, relationships and evolutionary dynamics of invertebrate animals over Earth's history, particularly over the Phanerozoic (i.e., the last 540 million years). Emphasis on the use of invertebrate fossil record to test ideas about long term evolutionary patterns as well as learning the histories and basic anatomies of major invertebrate taxa.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:SPRING

Credit Hours:3

ACE:

BIOS852
Field EpidemiologyCrosslisted with BIOS 452

Prerequisites: LIFE 121; LIFE 121L; three hours of BIOS

Offered summers only at Cedar Point Biological Station.

Description: Principles of epidemiology and the role in modern medicine. Combination of theory and practice with living populations.

This course is a prerequisite for: VBMS 949

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Offered:SUMMER
Experiential Learning:Fieldwork

Credit Hours:4

ACE:

BIOS853
Predator EcologyCrosslisted with BIOS 453

Prerequisites: BIOS 207 or NRES 220

Description: Conservation and management of native and invasive predators. Functional and numerical responses. Evolution of predator-prey interactions. Optimal foraging. Modeling predator-prey population dynamics. Trophic cascades. Prey defenses against predation.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Offered:SUMMER
Experiential Learning:Fieldwork

Credit Hours:4

ACE:

BIOS854
Ecological InteractionsCrosslisted with BIOS 454, NRES 454, NRES 854

Prerequisites: LIFE 121; LIFE 121L; BIOS 207 or NRES 220; Senior Standing

Description: Nature and characteristics of populations and communities. Interactions within and between populations in community structure and dynamics. Direct and indirect interactions and ecological processes, competition, predation, parasitism, herbivory, and pollination. Structure, functioning and persistence of natural communities, foodweb dynamics, succession, and biodiversity.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Graded

Credit Hours:3

ACE:

BIOS856
Mathematical Models in BiologyCrosslisted with BIOS 456, NRES 456, NRES 856

Description: Biological systems, from molecules to ecosystems, are analyzed using mathematical techniques. Strengths and weaknesses of mathematical approaches to biological questions. Brief review of college level math; introduction to modeling; oscillating systems in biology; randomness in biology; review of historically important and currently popular models in biology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS857
Ecosystem EcologyCrosslisted with BIOS 457, GEOL 457, GEOL 857

Prerequisites: BIOS 207 and CHEM 110A and 110L and Senior standing

Description: Processes controlling the cycling of energy and elements in ecosystems and how both plant and animal species influence them. Human-influenced global and local changes that alter these cycles and ecosystem functioning.

This course is a prerequisite for: BSEN 954, NRES 954

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option

Credit Hours:4

ACE:

BIOS859
LimnologyCrosslisted with BIOS 459, NRES 459, NRES 859

Description: Physical, chemical, and biological processes that occur in fresh water. Organisms occurring in fresh water and their ecology; biological productivity of water and its causative factors; eutroplication and its effects.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Offered:SPRING
Course and Laboratory Fee:$85

Credit Hours:4

ACE:

BIOS860
Soil Microbial EcologyCrosslisted with PLAS 460, BIOS 460, NRES 460, SOIL 460, AGRO 860, NRES 860

Prerequisites: Senior standing.

Recommend having a strong science background, including courses from the agronomic, environmental, microbiology, engineering or medicine disciplines.

Description: Soil from a microbe's perspective-growth, activity and survival strategies; principles governing methods to study microorganisms and biochemical processes in soil; mechanisms controlling organic matter cycling and stabilization with reference to C, N, S, and P; microbial interactions with plants and animals; and agronomic and environmental applications of soil microorganisms.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:SPRING

Credit Hours:3

ACE:

BIOS862
Animal BehaviorCrosslisted with BIOS 462

Prerequisites: BIOS 206, 207 and Senior Standing

Description: Introduction to animal behavior stressing the ethological approach. Anatomical and physiological bases of behavior, ontogenetic and phylogenetic observations, and the relations of animal behavior studies to genetics, ecology, taxonomy, and evolution.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS865
Behavioral NeuroscienceCrosslisted with BIOS 465, PSYC 465, PSYC 865

Prerequisites: PSYC 273

Description: Relationship of physiological variables to behavior, an introduction to laboratory techniques in neuropsychology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS866
Advanced Methods in NeurobiologyCrosslisted with BIOS 466

Prerequisites: LIFE 120

BIOS 206 and BIOS 213 are recommended.

Description: Examination of current technologies and methodologies used in neuroscience pertaining to medical, laboratory, and industrial applications across several organisms.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:SPRING

Credit Hours:3

ACE:

BIOS871
Plant SystematicsCrosslisted with BIOS 471

Prerequisites: LIFE 121 and LIFE 121L.

Description: Overview of the diversity of plants and algae, with emphasis on phylogenetic relationships, the evolution of important physical and genomic characteristics, principles of plant classification and identification, and modern methods of plant molecular systematics. Lab work on taxonomic analysis and plant identification.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$15

Credit Hours:4

ACE:

BIOS874
HerpetologyCrosslisted with BIOS 474, NRES 474, NRES 874

Prerequisites: NRES/BIOS 386

Description: Fossil and living amphibians and reptiles. Anatomy, classification, ecology and evolution.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Offered:FALL
Course and Laboratory Fee:$50

Credit Hours:4

ACE:

BIOS875
Avian BiologyCrosslisted with BIOS 475, NRES 471, NRES 871

Prerequisites: LIFE 121 & LIFE 121L

May also be offered at Cedar Point Biological Station.

Description: Biology of birds emphasizing the behavior and ecology of this group. Topics include avian diversity, systematics & evolutionary history, flight, foraging, migration, communication, reproductive biology, population ecology and conservation biology.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Experiential Learning:Fieldwork

Credit Hours:3

ACE:

BIOS875L
Avian Biology LaboratoryCrosslisted with BIOS 475L

Prerequisites: Parallel registration in BIOS 475/875

Description: Avian field identification in diverse prairie, riparian, and montane habitats. Individual studies of foraging behavior, territoriality, anti-predator behavior, mating systems, or nesting ecology.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Grading Option:Grade Pass/No Pass Option

Credit Hours:1

ACE:

BIOS876
MammalogyCrosslisted with BIOS 476, NRES 476, NRES 876

Prerequisites: 8 hrs BIOS; BIOS/NRES 386 or NRES 311.

May also be offered at Cedar Point Biological Station. Field trips are required and may occur outside of scheduled class time. Lab and field time emphasize diversity of mammalian families and species identification of Nebraska mammals.

Description: Evolution, natural history, ecology, and functional morphology of planetary mammals and mammals of the Northern Great Plains.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$25

Credit Hours:4

ACE:

BIOS877
Bioinformatics and Molecular EvolutionCrosslisted with BIOS 477

Prerequisites: BIOS 206 or parallel; CHEM 251 or CHEM 255 or CHEM 261.

Statistics course recommended.

Description: Pairwise and multiple alignments, sequence similarity and domain search, distance estimation, phylogenetic methods, gene mining, protein classification and structure. Algorithms used in bioinformatics as well as fundamental concepts of molecular evolution that underlie various bioinformatics methods.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS878
Plant AnatomyCrosslisted with BIOS 478, PLAS 478, AGRO 878, HORT 878

Prerequisites: 8 hrs biological sciences

Description: Development, structure, and function of tissues and organs of the higher plants. Relationships of structure to physiology and ecology of plants.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option

Credit Hours:4

ACE:

BIOS880
Ecology and Evolution of ArachnidsCrosslisted with BIOS 480

Prerequisites: BIOS 207 or NRES 220

Description: Ecology and evolutionary biology of living arachnids.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option

Credit Hours:4

ACE:

BIOS885
Aquatic InsectsCrosslisted with BIOS 485, ENTO 402, ENTO 802, NRES 402, NRES 802

Prerequisites: 12 hrs biological sciences.

Description: Biology and ecology of aquatic insects.

This course is a prerequisite for: BIOS 485L, BIOS 885L, ENTO 402L, ENTO 802L, NRES 402L, NRES 802L

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Grading Option:Grade Pass/No Pass Option

Credit Hours:2

ACE:

BIOS885L
Identification of Aquatic InsectsCrosslisted with BIOS 485L, ENTO 402L, ENTO 802L, NRES 402L, NRES 802L

Prerequisites: Parallel ENTO 802, NRES 402/802, BIOS 485/885.

Description: Identification of aquatic insects to the family level.

Course details
Credit Hours:1
Max credits per semester:1
Max credits per degree:1
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$25

Credit Hours:1

ACE:

BIOS887
Field ParasitologyCrosslisted with BIOS 487

BIOS 207 or NRES 220 recommended. Offered summers only at Cedar Point Biological Station.

Description: Animal host-parasite relationships, epizootiology, ecology, host distribution, classification, and life cycle stages of animal parasites.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Experiential Learning:Fieldwork

Credit Hours:4

ACE:

BIOS889
IchthyologyCrosslisted with BIOS 489, NRES 489, NRES 889

Prerequisites: LIFE 120 and LIFE 121

May also be offered at Cedar Point Biological Station.

Description: Fishes, their taxonomy, physiology, behavior, and ecology. Dynamics of fish stocks and factors regulating their production.

Course details
Credit Hours:4
Max credits per semester:4
Max credits per degree:4
Grading Option:Grade Pass/No Pass Option
Course and Laboratory Fee:$20
Experiential Learning:Research

Credit Hours:4

ACE:

BIOS891
Special Topics in Biological SciencesCrosslisted with BIOS 491

Prerequisites: BIOS 206 or BIOS 207

Description: Topics vary.

Course details
Credit Hours:1-4
Max credits per semester:4
Max credits per degree:9
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-4

ACE:

BIOS899
Masters Thesis

Prerequisites: Admission to masters degree program and permission of major adviser

Course details
Credit Hours:1-10
Max credits per semester:10
Max credits per degree:99
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-10

ACE:

BIOS915
Graduate Seminar

Prerequisites: Permission

Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS915A
Genetics, Cellular and Molecular Biology (GCMB)
Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS915B
Graduate Seminar - Behavioral Ecology
Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS915E
Graduate Seminar - Ecology, Evolution and Behavior (EEBE)
Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS915N
Population Ecology
Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS915P
Graduate Seminar - Parasitology
Course details
Credit Hours:1-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-3

ACE:

BIOS920
Viral Evolution

Description: The mechanisms by which DNA and RNA viruses evolve. The relationships between virulence, attenuation and host selection. Transfer of genetic material between virus and host. Evolution of HIV leading to escape from immunologic pressure or drug selection.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Graded

Credit Hours:3

ACE:

BIOS922
Viral Oncology

Description: Viral oncogenes and the cellular pathways that they influence. The mechanisms by which DNA and RNA viruses cause cancer. Discovery of novel tumor viruses.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Graded

Credit Hours:3

ACE:

BIOS932
ProteinsCrosslisted with BIOC 932, CHEM 932

Prerequisites: BIOC/BIOS/CHEM 832 or BIOC/BIOS/CHEM 839

Description: Protein structure and function.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Grading Option:Grade Pass/No Pass Option

Credit Hours:2

ACE:

BIOS933
EnzymesCrosslisted with BIOC 933, CHEM 933

Prerequisites: BIOC/BIOS/CHEM 432/832, or BIOC/BIOS/CHEM 839

Description: Kinetics regulation and reaction mechanisms of enzymes.

Course details
Credit Hours:2
Max credits per semester:2
Max credits per degree:2
Grading Option:Grade Pass/No Pass Option

Credit Hours:2

ACE:

BIOS934
Genome Dynamics and Gene ExpressionCrosslisted with BIOC 934, CHEM 934

Prerequisites: BIOC/BIOS/CHEM 832

Description: Detailed examination of dynamic control mechanisms of genome maintenance and gene regulation. Mechanisms of transcription, translation, and replication based on analysis of current and seminal literature.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS935
Metabolic Function and DysfunctionCrosslisted with BIOC 935, CHEM 935

Prerequisites: BIOC/CHEM/BIOS 432/832 and permission

Description: Current metabolic research at the bioenergetic, metabolomic, and molecular level. The normal metabolic processes that go awry in cancer, obesity, and oxidative stress.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS942
Genetics, Genomics, and Bioinformatics of ProkaryotesCrosslisted with VBMS 942

Description: Prokaryotic gene regulation, DNA exchange, DNA recombination and repair, comparative prokaryotic genomics and computer-based methods of analysis.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS945
RNA Biology

Prerequisites: BIOS 820

Description: Role of RNA in regulation of gene expression and in determining genome structure. Regulation of mRNA stability and function, RNA as regulatory molecules and enzymes, and computer-based methods of analysis.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS947
Industrial Microbiology and Biotechnology

BIOS 840 recommended.

Description: Biosynthetic activity of bacteria, yeasts, and fungi, including genetically engineered organisms: antibiotic, amino acid, enzyme, and vitamin production; polysaccharides, steroid transformation, microbes as food sources, microbial insecticides, petroleum microbiology, fermentation engineering, and mass production of microbial cells.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS952
Likelihood & Bayesian Ecology

NRES/STAT 803 recommended.

Description: Covers the use of maximum likelihood and Bayesian analysis in analyzing ecological data broadly defined. A conceptual understanding of the statistical tools is emphasized as well as a practical experience of conducting the analysis using real data and current software.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option
Offered:FALL

Credit Hours:3

ACE:

BIOS960
Biosystematics and NomenclatureCrosslisted with ENTO 960

Description: Methods and principles of systematics and nomenclature.

Course details
Credit Hours:2-3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:2-3

ACE:

BIOS962
Animal Communication

Prerequisites: BIOS 804

Description: Course work in physics recommended. Advanced introduction to the evolution of animal communication. Addresses evolution of signal structure (including acoustic, visual, electrical, and chemical signals), environmental effects on signal transmission, and the evolution of receiver responses to signals.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS964
Signal TransductionCrosslisted with VBMS 964

Prerequisites: BIOS 820.

Description: Molecular basis of genetics in eukaryotes. Gene structure and regulation, transposable elements, chromosome structure, DNA replication and repair mechanisms and recombination.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS967
Introduction to R for Biological Sciences

Description: Gain practical knowledge of the R programming language for statistical analysis, graphics, data management and simulations in the biological sciences.

Course details
Credit Hours:3
Max credits per semester:3
Max credits per degree:3
Grading Option:Grade Pass/No Pass Option

Credit Hours:3

ACE:

BIOS996
Research

Prerequisites: Permission of instructor and departmental Graduate Committee

Description: Research other than thesis.

Course details
Credit Hours:3-10
Max credits per semester:10
Max credits per degree:10
Grading Option:Grade Pass/No Pass Option

Credit Hours:3-10

ACE:

BIOS998
Special Topics in the Life Sciences

Prerequisites: Permission

Description: Reviews of specialized subject areas. Subject dependent on student demand and availability of staff.

Course details
Credit Hours:1-24
Max credits per semester:24
Max credits per degree:24
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-24

ACE:

BIOS999
Doctoral Dissertation

Prerequisites: Admission to doctoral degree program and permission of supervisory committee chair

Course details
Credit Hours:1-24
Max credits per semester:24
Max credits per degree:99
Grading Option:Grade Pass/No Pass Option

Credit Hours:1-24

ACE: